SquirrelWeb has references for many publications from 1999 to present (by year and author). The Forestry Commission (FC) Research Information Service can be searched for FC information notes and other squirrel-related articles and publications.  (Note this is being setup for anyone to access (within and outside FC) from Feb 2013 so link is not available yet).

Below are the key publications within the following subjects:

Policy and legislation

Site and species management


Practical support of local Red squirrels

Red squirrel on feederRegional and local Red squirrel groups encourage people to get involved with their conservation work. If you are interested in helping, please follow the links for the area you are in.

The UKRSG does not generally recommend supplementary feeding for red squirrels (Please see advice note); However if red squirrels are already coming to your garden, or you wish to attract them please read the following UKRSG guidance. Do not feed if both red and grey squirrels are present.

  • Supplementary Feeding in Gardens (PDF-123K)
    UKRSG Advice Note C1, 2004.
  • Shuttleworth CM (1997). The effect of supplemental feeding on the diet population density and reproduction of red squirrels.
    In: The Conservation of Red Squirrels, Sciurus vulgaris L. (eds J. Gurnell & P.W.W. Lurz), pp.13-24 Peoples Trust for Endangered Species, London.
  • Shuttleworth C.M. (2001). Traffic related mortality in a red squirrel (Sciurus vulgaris) population receiving supplemental feeding.
    Urban Ecosystems, 5(2);109-118.

Squirrelpox virus and Adenovirus

  • Everest, D.J., Shuttleworth, C.M., Grierson., S.S., Duff, J.P., Jackson, N., Litherland, P., Kenward, R.E. & Stidworthy, M.F. (2012). Systematic assessment of the impact of adenovirus infection on a captive reintroduction project for red squirrels. Veterinary record. 100617.
  • McInnes, C. et al. (2012). The emergence of squirrelpox in Ireland; Animal Conservation.
  • Squirrel Poxvirus - general information (PDF-56K)
    UKRSG Advice Note E1, 2004 amended.
  • Thomas et al. (2003). A Novel Poxvirus Lethal to Red Squirrels (PDF-126K)
    Short communication. Journal of General Virology, 84: 3337-3341.
  • Duff et al. (2007). Enteric adenovirus infection in a red squirrel (Sciurus vulgaris).
    The Veterinary Record March 17, 2007: 384.
  • Rushton et al, (2006). Disease threats posed by alien species: the role of a poxvirus in the decline of the native red squirrel in Britain. Epidemiology and Infection 134: 521-533.
  • Tompkins, D., Sainsbury, A.W., Netteleton, P., Buxton, D. & Gurnell. J. (2002). Parapoxvirus causes a deleterious disease of red squirrels associated with UK population declines.
    Proceedings of the Royal Society of London 269: 529-533.

Squirrel ecology

  • Bosch, S. & Lurz, P. (2012). The Eurasian Red Squirrel. ISBN 978-3894322588.
  • Gurnell, J. (1987). The Natural History of Squirrels.
    London, Christopher Helm.
  • Gurnell, J. (1991). Grey squirrel. The Handbook of British Mammals.
    G. B. Corbet and S. Harris, Blackwell Scientific: 186-91.
  • Gurnell, J., Lurz, P.W.W. & Wauters, L.A. (2012). Squirrels. The Mammal Society, Southampton, UK. Pp. 35.
  • Harris, S. & Yalden, D.W. (2008). Mammals of the British Isles: Handbook, 4th Edition.
    The Mammal Society.
  • Koprowski, J. L. (1994). Sciurus carolinensis.
    Mammalian Species. 480: 1-9.
  • O'Teangana, D. (1999).
    The distribution and ecology of the red squirrel (Sciurus vulgaris) and grey squirrel (Sciurus carolinensis) in Northern Ireland.
    The Queens University of Belfast. Unpublished PhD thesis.
  • Lurz, P.W.W., Garson, P.J. & Rushton, S.P., (1995). The ecology of squirrels in spruce dominated plantations: implications for management. Forest Ecology and Management, 79, 79-90.
  • Tonkin, J. M. (1983). Ecology of the Red Squirrel (Sciurus vulgaris L.) in mixed woodland.
    University of Bradford.