Forestry Commission publications
Mayle, B., Webber, J. & Jones, B. (2012). Aerial surveys for P. ramorum in Wales show extent of damage by squirrels to larch. Ecotype Winter 2012.
Mayle, B., Ferryman, M., Pepper, H. (2007). Controlling Grey Squirrel Damage to Woodlands (PDF-1779K). Forestry Commission Practice Note 4 (Revised). Forestry Commission, Edinburgh.
Ferryman, M. and Mayle, B. (2005). Grey squirrel damage prediction - interim progress. Ecotype 30, 1-2.
Bryce, J., Cartmel, S. and Quine, C. P. (2005). Habitat use by red and grey squirrels: results of two recent studies and implications for management (PDF-546K). Forestry Commission Information Note 76. Forestry Commission, Edinburgh.
Mayle, B. A. (2004) Grey squirrel management in woodlands, in Managing woodlands and their mammals (Quine, C., Shore, R. & Trout, R., Eds.): proceedings of a symposium organised jointly by the Mammal Society and the Forestry Commission, 45-49, Forestry Commission, Edinburgh.
Gurnell, J., Lurz, P.W.W., Pepper, H. (2001) Practical techniques for surveying and monitoring squirrels (PDF-3830K), Forestry Commission Practice Note 11, Forestry Commission, Edinburgh.
Pepper, H., Bryce, J., Cartmel, S. (2001), Squirrel management: 2. Co-existence of red squirrels and grey squirrels, in Forest Research Annual Report and Accounts 1999-2000. Forestry Commission, Edinburgh.
Pepper, H., Moore, H. (2001) Squirrel management: 1. Development of a contraceptive vaccine for the grey squirrel, in Forest Research Annual Report and Accounts 1999-2000. Forestry Commission, Edinburgh.
Pepper, H., Patterson, G. (2001) Red squirrel conservation (PDF-250K), Forestry Commission Practice Note 5 (revised). Forestry Commission, Edinburgh.
Pepper, H. (1998) Nearest neighbour method for quantifying wildlife damage to trees in woodland (PDF-139K), Forestry Commission Practice Note 1. Forestry Commission, Edinburgh.
Mayle, B.A., Gurnell, J. (1991) Edge management and small mammals. in Edge management in woodlands (Ferris-Kaan, R., Ed.), Forestry Commission Occasional Paper 28. 432-449, Forestry Commission, Edinburgh.
Nichols, C.P., Gill, R.M.A., Drewe, J.A. Goode, N. Gregory, N. (2016) A novel causal mechanism for grey squirrel bark stripping: The Calcium hypothesis. Forest Ecology and Management. 367 12-20.
Mayle, B., Ferryman, M., Peace, A., Yoder, C., Miller, L., Cowan, D., (2013) The use of DiazaCon™ to limit fertility by reducing serum cholesterol in female grey squirrels, Sciurus carolinensis. Pest Management Science 69, 414-424.
Mayle, B.A. and Broome, A. (2013) Changes in the impact and control of an invasive alien, the grey squirrel Sciurus carolinensis, as determined from regional surveys. Pest Management Science, 69, 323-333.
Willoughby, I., Jinks, R.L., Morgan, G.M., Pepper, H., Budd, J. and Mayle, B.M. (2011). The use of repellents to reduce predation of tree seed by wood mice (Apodemus sylvaticus L.) and grey squirrels (Sciurus carolinensis Gmelin). European Journal of Forest Research. 130, 4, 601-611.
Yoder, C. A., Mayle, B.A., Furculow, C.A., Cowan, D,P. & Fagerstone, K.A. (2011). Feeding of grey squirrels (Sciurus carolinensis) with the contraceptive agent DiazaCon: effect on cholesterol, hematology, and blood chemistry. Integrative Zoology 6: 409-419.
Mayle, B. A., Proudfoot, J., Poole, J. (2009). Influence of tree size and dominance on incidence of bark stripping by grey squirrels to oak and impact on tree growth. Forestry 2009,82,4, 431-444. Full text, PDF (201K)
Ferryman, M., Mayle, B. A. and Morgan, G. W. (2006). Visual method for evaluating the state of sexual development in male grey squirrels (Sciurus carolinensis). Reproduction, Fertility and Development 18, 383-393.
Mayle, B. A. (2005). Britain’s woodlands under threat; Grey squirrels and the risk they pose to European woodlands. Trees, Journal of the International Tree Foundation. 65 9-11.
Pepper, H.W., G. Kerr (2005). Preliminary studies on collars to protect trees from grey squirrel bark-stripping damage. Quarterly Journal of Forestry 99(2): 105-112.
Gurnell, J., Lurz, P.W.W., Shirley, M.D.F., Cartmel, S., Garson, P.J., Magris, L., Steele, J. (2004) Monitoring red squirrels (Sciurus vulgaris) and grey squirrels (Sciurus carolinensis) in Britain (PDF-347K), Mammal Review. 34, 51-74.
Hewson, C., Fuller, R., Mayle, B., Smith, K. (2004) Possible impacts of grey squirrels on birds and other wildlife. British Wildlife 15, 183-191.
Mayle, B., Gurnell, J. (2004) Squirrel control in conifers, Forestry and British Timber, April 2004, 16-19.
Gurnell, J., Mayle, B.A. (2003) Ecological impacts of the alien grey squirrel (Sciurus carolinensis) in Britain, conference proceedings: MammAliens - a one day conference on the problems caused by non-native British mammals, 40-45, Mammals Trust UK.
Gurnell, J. (1999) Grey squirrels in woodlands: managing grey squirrels to prevent woodland damage. Enact 7, 10-14.